The Ants of Africa New records of ants from Sudan Acacia senegal collected by J H Mathews, 2000-1

Ants from Sudan Acacia senegal, collected by J H Mathews, University of Oxford, 2000-1

* first record for Sudan

Collection localities (P = Plantation, SNW = semi-natural woodland); ants were not captured from all plots

Preliminary trials were in Kordofan, El Ain, near El Obeid, in October 2000; location not given but presumably close to the Rahad plantation (examined 040, 059, 072-096)

Rahad Research Area, Kordofan - 12°56' N 30°35' E; Nov-Dec 2000 - 15 P 12 SNW; June-July 2001 - 21 P 18 SNW (examined 4 tubes; 273-281); Sept-Oct 2001 - 21 P 20 SNW (examined 9 tubes; 310-340)

Khordonia, Damazine Plantation, Blue Nile - 11°51' N 34°15' E; Nov-Dec 2000, 17 P 29 SNW; June-July 2001 - 12 P 20 SNW (examined 35 tubes from 222-297); Sept-Oct 2001 - 21 P, 20 SNW (examined 34 tubes from 341-381 & 421-423)

Abu Gmein, Blue Nile - 11°21' N 34°27' E; Nov-Dec 2000 - 21 P 20 SNW (examined 12 tubes; 115-172); June-July 2001 - 21 P 21 SNW (examined 7 tubes; 174-217); Sept-Oct 2001 - 20 P 20 SNW (examined 24 tubes; 383-414)

Not seen by BT - Khordonia Nov-Dec 2000

Mathews (thesis, Appendix D, pp 327-8) separated 41 Formicidae RTUs.  Cites 340 samples; with each canopy sample given a single number, although split into taxonomic groups. [BT comments - almost irreconcilable; clearly I have not had all the samples; the Kordofan findings are not in the thesis]

DOLICHODERINAE
Tapinoma sp 02 (12) [unclear as I have separated two species, from 5 trees]
DORYLINAE
Dorylus sp 01 (Khordonia SNW, 1) [a misidentification as none in the tubes and Dorylus species very rarely, if ever ascend trees]
FORMICINAE
Anoplolepis sp 01 (7) [possibly a misidentification of Lepisiota gracilicornis as Anoplolepis gracilipes]
Camponotus sp 01 (19); Camponotus sp 02 (Rahad P, 1); Camponotus sp 03 (Abu Gmein, 1); Camponotus sp 04 (18); Camponotus sp 06 (Abu Gmein 1) [the only species I found was Camponotus sericeus although there are variations in the colour, notably in the reddishness of the head]
Lepisiota sp 01 (29 trees, common)
Lepisiota sp 02 (Rahad SNW, 2 trees, 10)
Lepisiota sp 03 (Khordonia 1, Rahad 1)
Lepisiota sp 04 (Abu Gmein 1, Rahad 1)
Lepisiota sp 06 (145 trees, very common)
Plagiolepis sp 01 (Abu Gmein p, 1 tree, 1; Khordonia SNW, 1 tree, 1; Rahad SNW, 2 tree, 3) [
Plagiolepis sp 02 (28) [probably Plagiolepis sudanica and Plagiolepis brunni combined]
MYRMICINAE
Cardiocondyla sp 01 (Khordonia P, 2 trees, 6; Khordonia SNW, 2 trees, 11; Abu Gmein SNW, 1 tree, 2) [only specimen seen was a single queen]
Crematogaster sp 01 (Rahad SNW, 1 only) [don't know]
Crematogaster sp 02 (23) [probably a combination of the smaller Crematogaster (Cr.  spp, most being Cr. acaciae]
Crematogaster sp 03 (Khordonia P, 1 tree, 3) [possibly the Crematogaster (Cr.) senegalensis Eritrea form]
Diplomorium sp 1 (Rahad SNW, 1) [possibly a major worker of Monomorium oscaris]
Eutetramorium sp 01 (Rahad SNW (June-July 2001), 1 tree, 123) [definite mis-identification but possibly not among the tubes I have seen]
Myrmicinae indet 01 (Rahad SNW, 1)
Leptothorax sp 01 (10 trees, 37) [possibly Tetramorium cristatum]
Leptothorax sp 02 (Rahad P 1, Abu Gmein P, 4 trees 8) [probably Nesomyrmex angulatus]
Monomorium sp 01 (10 trees, 36)
Monomorium sp 02 (Rahad, 3 trees, 9)
Monomorium sp 03 (Abu Gmein 2 trees, Khordonia 2 trees,5)
Monomorium sp 04 (15 trees, up to 20)
Monomorium sp 05 (Rahad SNW, 2 trees, 8)
Oligomyrmex sp 01 (Khordonia SNW 1 tree, 2) [possibly minors workers of Monomorium oscaris]
Pheidole sp 01 (Khordonia 3 trees, 8; Rahad, 1 tree, 5)
Pheidole sp 02 (Khordonia P, 1)
Pheidole sp 03 (Rahad SNW 1 tree, 2)
Pheidole sp 04 (Rahad P, 1) [not in tubes I have had]
Pristomyrmex n sp 01 (Abu Gmein, 2 trees P, 13; Khordonia P, 1 tree, 3; Khordonia SNW, 1 tree, 3) [definite misidentifications; I found an example of Plagiolepis brunni labelled as "Pristomyrmex"]
indet sp 01 (Abu Gmein P 1)
indet sp 02 (Khordonia P 1)
indet sp 03 (Abu Gmein SNW 1)
indet sp 04 (Khordonia SNW 1 tree, 3)
indet sp 05 (Abu Gmein P, 1 tree, 2; Rahad SNW, 1)

Species List - 48 species

SUBFAMILY	Species	Image
DOLICHODERINAE	Tapinoma carininotum Weber type location Sudan Kordofan, JM 091, 1 tree (1) Abu Gmein, JM 402, 1 tree (1) Probably JM Tapinoma sp 02, combined with T. luteum Mounted and photographed JM 091 and JM 402
0DOLICHODERINAE	Tapinoma luteum (Emery) JM 330 east and southern Africa * Kordofan 2 trees (2 & 5) Rahad 2 trees (4 & 8) Khordonia 1 tree (1) Abu Gmein 1 tree (2) Mounted and photographed JM 092, JM 283, JM 330 and JM 334
FORMICINAE	Camponotus (Orthonotomyrmex) sericeus (Fabricius) JM 223, dimorphic pan-African, common Kordofan 8 trees Rahad 8 trees Khordonia 14 trees Abu Ghmein once Note there is a colour variation with some having a brighter red head - this may explain the JM separation into four species Mounted and photographed JM 223 and JM 336
FORMICIDAE	Lepisiota affinis (Santschi) black form JM 406 (Abu Gmein) Head sub-square; clypeus with a strong median carina/keel; head with a distinct median impressed line, reaching back to posterior level of the eyes; eyes ovoid but quite broad with angled forward edge; abundant long erect hairs (ca 0.17 mm); petiole profile moderately thick with obtuse apex, shorter than propodeal declivity TL ca 2.5 mm, CI 85, SI 127 Matches type description Mounted and photographed JM 341, JM 353, JM 406 and JM IT10
FORMICIDAE	Lepisiota affinis (Santschi) brown form JM 341 Kenya & Sudan Khordonia 4 trees (19) Abu Gmein 1 tree (1) Omer Sudan 2-2-1, Wad Medani
FORMICIDAE	Lepisiota sp JM 234 head sub-square; eyes relatively large (ca 0.40 X length of side of head) shallowly convex; petiole scale with a narrow profile, as high as the propodeum declivity, weakly bispinose; erect pilosity moderately abundant but distinctly shorter (ca 0.10 mm) than on albata and canescens TL ca 2.6 mm, CI 89, SI 133 Khordonia 1 tree (numerous); label in tube JM "Lepisiota No 3" The description of Lepisiota imperfecta (Santschi), type location Zimbabwe is a close match, notably the relatively short sparser erect hairs and much reduced pubescence, but that had a TL 2.0 mm and was black Mounted and photographed JM 234
FORMICIDAE	Lepisiota albata (Santschi) Note polychromic from dark brown-black to dull yellow-brown; JM 372 longer erect hairs (ca 0.12 mm) than JM 234; very thin petiole scale (thinner than Santschi indicates) but squarish head shape matches; eyes relatively large (ca 0.38 X length of side of head) shallowly convex; TL ca 2.6 mm, CI 85, SI 130 Type location Zaire but I have others from Sudan; slightly larger than the type size, TL 2.3 mm. This commonly has colonies containing pale to near albino specimens, contrasting with the dark red-brown to black majority Kordofan 1 tree Rahad 1 tree Khordonia 33 trees Abu Gmein 31 trees Mounted and photographed JM 342, JM 343, JM 345, JM 372, JM 381, JM 400 and JM 422
FORMICIDAE	Lepisiota albata colour variation
FORMICIDAE	Lepisiota canescens (Emery) JM 402; head sub-rectangular; eyes ovoid and flat (ca 0.33 of length of side of head); quite thick petiole scale, with a barely impressed upper margin; erect hairs yellowish (ca 0.16 mm long); almost completely dark brown-black, tarsi and base of scapes dull yellow-brown TL ca 2.5 mm, CI 77, SI 135 common across Sahel zone Khordonia 3 trees (7) Abu Gmein 4 trees (9); 211 had a label in tube JM "Lepisiota No 3" Mounted and photographed JM 211, JM 402, JM 406 and JM 421 Omer Sudan 2009.11; Ar Roseris, 11°50'60"N 34°22'60"E Several historic records; Kaka, collector Jaegerskioeld (Mayr, 1904b), on acacia stem; Khartoum, Faschoda, Renk & Gondokoro, collector P Magretti (Mayr, 1907a); Khratoum, collector V Karavaiev (Karavaiev, 1911); Sennaar, El Obeid, Karshawal, collectors H Werner & R Ebner (Viehmeyer, 1923)
FORMICIDAE	Lepisiota curta (Emery) Head somewhat wider and more truncate than depilis, with scape surpassing occiput by < one-third of own length; eyes quite small TL 2.3-2.5 mm, CI 88, SI 103 but 117-124 on fresh Sudan specimens Mounted and photographed JM 370 Omer Sudan 11; Umdaldal 1, South Sudan Province, collector Awatif Omer, 2006 Omer Sudan 2.7; Dinder (Aldinder) National Park Centre, 11°21' N 35°02' E along Sudan-Ethiopian Boundary Omer Sudan 2009.7; Sennar, 13°33'N 033°37'E, collector Awatif Omer Omer Sudan 2009.26; Abbasiya, 12°1'9.01" N 27°58'3.17" E
FORMICIDAE	Lepisiota depilis (Emery) acholli (Weber) JM 351; clypeus with distinctive raised median carina; eyes near flat (ca 0.4 x length of side of head); alitrunk relatively short and stocky, pronotum quite strongly domed; petiole scale weakly bispinose, head narrowed anteriorly but in profile convex on both upper and lower sides; erect hairs moderately long and slender off-white (ca 0.10 mm long). head quite shaggy due to decumbent pubescence; overall uniform dark brown-black type location Somalia; type description has petiole scale effectively unarmed, head, pronotum and mesonotum subopaque. This is shinier but seems a good match for acholli TL ca 2.4 mm, CI 88, SI 124 Khordonia 2 trees (351 only 12; 370 3 plus very many others) Mounted and photographed JM 351
FORMICIDAE	Lepisiota foreli (Arnold) JM 073; head near rectangular, with a narrow profile; clypeus with an obtuse median carina; eye ovoid weakly convex (0.37 X length of side of head); petiole scale profile quite narrow, with a sharp apex, very weakly bispinose, unusually narrow seen from above; mahogany, almost no erect hairs except a few very short on gaster TL ca 2.2 mm, CI 82, SI 111 type location Zimbabwe * May not be foreli but generally matches description, small size and colour, here somewhat darker red-brown, alitrunk narrow in dorsal view; limited numbers of pale specimens (callows?), distinctive dull appearance of propodeum Kordofan 6 trees most with 1 ant but one tree (089) with numerous Khordonia 1 tree (1) Mounted and photographed JM 040, JM 073, JM 074, JM 080, JM 084, JM 087, JM 091 and JM 096
FORMICIDAE	Lepisiota gracilicornis (Forel) JM 040; slender. elongated species, petiole and propodeum with sharp spines TL ca 3.6 mm, CI 75, SI 230 Unmistakeable Horn of Africa species Kordofan 1 tree (1) Rahad 1 tree (1) Mounted and photographed JM 040 and JM IT6 Omer Sudan 1.16, Port Sudan, 19.57° N 37.27° E Omer Sudan 2.2.1, Wad Medani Omer Sudan 2009.17; Afad (South Ad Damazin), 11°46'0"N 34°21'0"E Omer Sudan 2009. 15; Shendi, Northern Sudan, 16°41'N 33°26'E Omer Sudan 2009.20.2 & 2009.21.1; Khartoum - 15°35'703" 32°32'15" E Omer Sudan 2009.28; Wad el Hadad - 13°49'12.06" N 33°32'31.53" E
FORMICIDAE	Lepisiota laevis (Santschi) JM 294; head sub-circular, widest at occipitum; eyes in full face view a peculiar shape, weakly convex (0.34 X length of side of head); short alitrunk and scapes; alitrunk strangely small, short and stocky; petiole scale weakly bispinose. also extremely narrow and small; shiny but head very finely sculptured and submatt; small, very sparse, short hairs (ca 0.07 mm long) on border of gaster segments; very dark brown * TL ca 2.4 mm, CI 89, SI 113 Khordonia, JM 294, 1 tree (1), JM 295 (1 worker) Kordofan, JM 092. JM 093 Mounted and photographed JM 092, JM 093, JM 294 and JM 295
FORMICIDAE	Lepisiota palpalis (Santschi) JM 174; small, long palps distinctive; head in full face view near circular; eyes large (0.43 x length of side of head); pronotum dorsally flattened; propodeum dorsum with a weak longitudinal septum; petiole bispinose; abundant fine erect yellow pilosity (on gaster ca 0.12 mm long); shiny unsculptured, except lateral mesonotum and propodeum which are very finely spiculate; chestnut to dark brown; antennae wholly yellow brown except for apical segments of funiculus TL ca 2.5 mm, CI 80, SI 138 Abu Gmein, JM 174 1 tree (5) - rather more brown than others from Sudan. Mounted and photographed JM 174 and JM 154 Also Omer 2009-34-2 Omer Sudan 20-01; Kosti, western bank of River Nile, 280 km south of Khartoum, collected by Awatif Omer, 2006
FORMICIDAE	Lepisiota palpalis Santschi type form JM 154; head subsquare, with weakly concave occiput; clypeus with quite a strong median carina/keel; eyes large (0.42 X length of side of head) ovoid but shallowly convex; petiole profile moderately narrowed but sharply tapered apically, bispinose; legs unusually long; abundant erect yellowish pilosity (ca 0.12 mm long); shiny but mesonotum and propodeum dull due to fine spiculation; dark brown TL ca 2.9 mm, CI 91, SI 124 Khordonia 1 tree (1) Mounted and photographed JM 154
FORMICIDAE	Lepisiota n sp JM IT6 Head sub-circular, in profile also sub-circular; frons with a long median impression; eyes moderate and set more or less on the front of the face, so that hardly breaking outline of sides (0.328 X length of side of head); scape unusually thick and apically widening; petiole scale with quite narrow profile, as high as the propodeal declivity; erect hairs abundant and quite thick (ca 0.12 mm long); base colour dusky yellow to dull yellow-grey, appearance overall is dull TL ca 2.6 mm, CI 89; SI 128 Khordonia 1 tree (1) tube with JM label "Lepisiota 4" Possibly the Santschi (1937d) "Acantholepis capensis Mayr v. subopaciceps n. v." (presently under L. simplex). Mounted and photographed JM IT6
FORMICIDAE	Lepisiota n sp JM 362 Very distinctive shiny unsculptured, yellow brown, gaster darker; pronotum dorsally flattened; head sub-square, widest behind the eyes; eyes ovoid moderately convex (0.37 X length of side of head); in profile head sub-circular with broadly convex lower side; clypeus with strong median carina running right to anterior margin; propodeum dorsum with distinctive longitudinal septum; petiole narrow and emarginate apically; abundant erect yellow pilosity (ca 0.16 mm long) TL ca 2.7 mm, CI 90, SI 131 Khordonia 1 tree (6) Mounted and photographed JM 362
FORMICIDAE	Lepisiota n sp JM 074 Small, occiput near straight; eyes almost flat and slightly inset into surface of head, narrow ovoid ca 1.7 longer than wide (0.37 X length of side of head); pronotum shallowly convex; petiole emarginate, moderately thick; sparse short yellow pilosity (ca 0.05 mm); shiny unsculptured, alitrunk yellow brown, head and gaster darker; TL ca 1.7 mm, CI 83, SI 130 Kordofan 1 tree (2) Mounted and photographed JM 074
FORMICIDAE	Lepisiota n sp JM 084 (also JM 072) Head sub-circular; eye relatively small, subcircular and flat (0.30 X length of side of head); petiole scale transversely unusually narrow, apex weakly concave, unusual in being slightly higher than the propodeal declivity; erect pilosity very much reduced and very short, ca 0.05 mm; weakly shiny and dull yellow-brown In curta-group of Santschi (1937d), with laevis (see above) TL ca 2.2 mm, CI 85-88, SI 113 Kordofan 2 trees (084 1 & 094 2) Mounted and photographed JM 072 and JM 084
FORMICINAE	Plagiolepis (Plagiolepis) brunni Mayr JM 115 Pan-African Khordonia 4 trees (7) [JM "Pristomyrmex"] Abu Gmein 6 trees (12) Mounted and photographed JM 115, JM 154, JM 183 JM 234, JM 385 and JM 398
FORMICINAE	Plagiolepis (Plagiolepis) exigua Forel (not sudanica) JM 160 matches type description although seemingly slightly lighter Kordofan 1 tree (19) Khordonia 2 trees (4) Abu Gmein 10 trees (12) Mounted and photographed JM 160, JM 172, JM 174, JM 211, JM 341, JM 413 and JM 414
FORMICINAE	Tapinolepis (Mesanoplolepis) mediterranea (Mayr) JM 072-01 major type location and only literature record was Egypt; Antweb has a photo of a specimen from Kenya * Kordofan 10 trees (1-20) Khordonia 4 trees (4) JM 289 had a label "Formicinae Plagiolepis" JM 279 had a label "Lepisiota No 4" Mounted and photographed JM 072, JM 092,  JM 279 and JM 289
FORMICINAE	Tapinolepis mediterranea JM 072-01 minor First illustration of minor
FORMICINAE	Tapinolepis (Mesanoplolepis) pernix (Viehmeyer) JM 092 Type location Sudan; matches type description; first illustrations Kordofan 7 trees (1-7) Khordonia 2 trees (2) JM 292 had label "Formicine Acropyga" Mounted and photographed JM 072 and JM 092
MYRMICINAE	Cardiocondyla emeryi Forel, queen JM 115 Pan-African and tramp species Abu Gmein 1 tree (1) Mounted and photographed JM 115
MYRMICINAE	Crematogaster (Crematogaster) acaciae Forel major JM 287 Type location Ethiopia from acacia * Kordofan 3 trees (8-numerous) Khordonia 5 trees (4-numerous) Mounted and photographed JM 092 and JM 287
MYRMICINAE	Crematogaster acaciae minor JM 287
MYRMICINAE	Crematogaster (Crematogaster) castanea F. Smith JM 174 poorly defined and very "variable" species; this matches var. ulugurensis Forel, from Tanzania, finely but densely striate head and alitrunk Khordonia 1 tree (1) Abu Gmein 1 tree (8) Mounted and photographed JM 174 and JM 370
MYRMICINAE	Crematogaster (Crematogaster) chiarinii Emery cincta Emery JM 294 The "sub-species" or variety was from Somalia; probably a distinct species as the type form is black and the variety was described as having a brown head, somewhat ferruginous in front, the alitrunk and pedicel ferruginous, the antennae and legs ferruginous-testaceous; the gaster with the base testaceous-yellow and the rest dull brown Khordonia 1 tree (1) Mounted and photographed JM 295
MYRMICINAE	Crematogaster (Crematogaster) mimosae Santschi JM 172 Type location Kenya, an Acacia inhabitant Abu Gmein 1 tree (150 +) Mounted and photographed JM 172
MYRMICINAE	Crematogaster (Crematogaster) rivai Emery major worker JM 072 Type location Ganana (Somalia/Ethiopia) from a swollen Acacia spine; close to type description, perhaps somewhat darker but with a distinctive 4-segmented antennal club; scapes thicker than other species * Kordofan 4 trees Mounted and photographed JM 072, JM 078, JM 091 and JM 336
MYRMICINAE	Crematogaster rivai minima worker
MYRMICINAE	Crematogaster (Crematogaster) senegalensis Roger JM 059 Pan-Sahelian species; this matches a type form specimen I have from Niger Kordofan 8 trees (2 trees have numerous workers) Mounted and photographed JM 040, JM 059, JM 084, JM 085 and JMN 294
MYRMICINAE	Crematogaster (Sphaerocrema) chlorotica Emery JM 174 Literature records are from West Africa but this matches the type description and specimens I have  from Mali and Benin Abu Gmein 1 tree (1) Mounted and photographed JM 174
MYRMICINAE	Monomorium nsp JM 076 JM 076 Distinctive striated mandibles Kordofan 1 tree (1 only); 24.x.2000 TL ca 3.4 mm, HW 0.71, HL 0.88, CI 90, SL 0.72, SI 102, PW 0.45; eyes ca 0.29 X HW Mounted and photographed JM 076
MYRMICINAE	Monomorium manir Bolton JM 254 Previously known only from the holotype collection in Kenya; from Acacia-Commophila scrub * first illustration Kordonia 1 tree (1) Mounted and photographed JM 254
MYRMICINAE	Monomorium mictilis Forel JM 174 Pan-Sahel species * Khordonia 2 trees (5 & 2) Abu Gmein 2 trees (1 & 1) Mounted and photographed JM 174, JM 345 and JM 394
MYRMICINAE	Monomorium niloticum Emery JM 183 Seemingly primarily North African and up the Nile to Sudan (my records) Khordonia 1 tree (1) Abu Gmein 2 trees (2 & 1) Mounted and photographed JM 183 and JM 244
MYRMICINAE	Monomorium oscaris Forel JM 093, polymorphic, major shown Pan-African savannah species Kordofan 1 trees (1 & several) Abu Gmein 2 trees (6 & 2) Mounted and photographed JM 040, JM 093 and JM 394
MYRMICINAE	Monomorium oscaris Forel JM 040, minor
MYRMICINAE	Monomorium pullulum Santschi JM 093, polymorphic, major shown Pan-Sahel, type location Senegal, but I have a specimen from Egypt, now these * Kordofan 10 trees (ca 12 on each) Rahad 1 tree (5) Khordonia 2 trees (many & 6) Mounted and photographed JM 072, JM 073 and JM 283
MYRMICINAE	Monomorium robustior Forel JM 093, polymorphic, major shown East African, type location Somalia * Kordofan 1 tree (1) Mounted and photographed JM 072
MYRMICINAE	Monomorium salomonis (Linnaeus) JM 040 Essentially a North African species Kordofan 1 tree (2) Mounted and photographed JM 040
MYRMICINAE	Myrmicine queen possibly Monomorium pullulum Santschi JM IT7 Rahad Plantation 1 tree (1) Mounted and photographed JM IT7
MYRMICINAE	Nesomyrmex angulatus (Mayr) JM 174 North Africa - Sahel species matches Egyptian specimens Abu Gmein 3 trees (2, 5 & 8) Mounted and photographed JM 174
MYRMICINAE	Pheidole escherichii Forel JM 334; minor type location Eritrea but I have specimens from Sudan Rahad 1 tree (1) Mounted and photographed JM 334
MYRMICINAE	Pheidole cf prelli JM 226; minor The type location for Pheidole prelli Forel is Kenya; and this is generally darker than the type description of overall yellow-rust * Khordonia 1 tree (1) Mounted and photographed JM 226
MYRMICINAE	Pheidole rugaticeps Emery JM IT2; minor type location Eritrea, but a Sahel species Khordonia 1 tree (1) Abu Gumein 1 tree (2) Mounted and photographed JM IT2
MYRMICINAE	Pheidole saxicola Wheeler JM 340; minor type location Zaire, I have a specimen from Congo * Rahad 1 tree (1) Mounted and photographed JM 340
MYRMICINAE	Pheidole sculpturata Mayr JM 240; minor type location South Africa, but recorded from Somalia and Kenya * Khordonia 1 tree (2) (first photo) Mounted and photographed JM 240
MYRMICINAE	Tetramorium cristatum Stitz JM 384 Seemingly a savannah species from right across the Sahel; with relatively few records (first photo) Abu Gmein 5 trees (3, 5, 5, 1 & 1) Mounted and photographed JM 384, Jm 385 and JM 390

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